✖Canon History
An insanely long time ago, a giant robot god made Autobots and Decepticons. They fought an epic and cheaply animated war on Earth, and the Autobots won. After a while they ran out of fuel or something (the writers are never really clear) and they all downgraded to smaller forms, as well as renaming the factions Maximals and Predacons, a strange choice of words since nobody had animal forms yet. The Predacons, descendants of the Decepticon losers, are a tiny portion of the population and treated as second class citizens. Still harbouring galaxy-conquering designs, they don't really appreciate this.
Pretty much nothing is known about Waspinator's life on his home planet, Cybertron. Some parts of fandom would peg him as a prostitute. This is a falsity. A huge, huge falsity. He was likely a petty thief or even an assembly line grunt. He probably didn't have an office job as he's too dumb to hold one down. What we do know is that he signed up for a mission led by Predacon renegade Megatron to steal an artifact, travel to Earth, and change the future or something. It's doubtful that he even understood the whole thing, he probably just liked the big shiny disc. Somehow, he was hired onto this mission as a pilot, a job usually associated with some kind of skill.
Chased to Earth by a bunch of Maximal scientists and their pet cheetah, the Predacons engaged in what was later to be called the Beast Wars and Fought The Good, Predaconial Fight. Meaning that they were a bunch of sneaky, underhanded bastards, and they still lost. Throughout this time, Waspinator was shot, crushed, or just plain blown up on pretty much every single mission. He was briefly possessed by the ghost of the Decepticon Starscream; sadly the only time that he showed any kind of intelligence or ambition.
Near the very end of the Beast Wars, right before everyone started dying, Waspinator declared that he was fed up. He was tired of following Megatron's orders, being the loyal and oh-so-expendable soldier. The good guys never blew up, right? So he'd become one of them! Then his teammates blew him to pieces and left him in the woods. By an odd twist of fate, this meant that he was out of harm's way for the final confrontation, making him pretty much the only Predacon to survive the Beast Wars. Karma is weird.
Finally, the Maximals found his scattered parts, put him back together out of pity, and left him alone on prehistoric Earth while they took Megatron back to Cybertron to stand trial. Alone with the neanderthals, who couldn't blow him up, Waspinator expressed that he was finally happy.
✖Cape & Cowl History
Waspinator arrived in the City dazed, confused, and slightly schizophrenic. Not only had the Porter given him a new, organic human body, but it had seen fit to return to his head the immortal spark of Starscream. Starscream spent most of his time berating Waspinator on his stupidity and frequent failures, taking over the body with little warning. Waspinator managed with this for a while, though Terrorsaur had to live across the hall from the multi-voiced crackpot and was not so complacent. Starscream harboured a tendeny to keep quiet around Megatron after his appearance, however. After this had been going on for a few months, Starscream's spark vanished. Waspinator was overjoyed, thinking that his luck had finally turned around. In reality, it was the arrival of another version of Starscream that made the spark disappear, this Starscream having memories of his first possession but no recollection of any time in the City. Shortly afterwards, Terrorsaur vanished from the City without a trace, in the way of those sent back to their homeworlds. Now Waspinator is left with a tyrannical boss, the lackey that was living in his mind, and no partner to put him back together when he gets shot. Such is life.
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